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Flashback: When Itachi met Shisui, one of his eyes were already stolen.Naruto Chapter 550 Title: “Koto Amatsukami”
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Itachi breaks free from edo tensei control and uses amaterasu on the summons. The crow's sharingan was Shisui's sharingan and Itachi uses shisui's jutsu to regain control.
Shisui's genjutsu allows the target to be unknowingly controlled and it is the strongest genjutsu. "Protect Konoha" was the order implanted into Shisui's eyes. The eye was then imlpanted to the crow and handed over to Naruto.
Itachi thought Sasuke would transplant his eyes and it should call out the crow when Sasuke activates his sharingan. Shisui's genjutsu would then activate and make Sasuke "protect konoha".
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